Hello,
Why do you mention mesh on the post title? It looks more like a star topology. Maybe doing a proper mesh solves your issue or doing a direct VPN to the desired site.
Lookin into your issue it seems like a domain encryption issue. I had a similar case a few years back, we had to add the performed NAT to the peer's encryption domain. Would like you to provide exact information but I can't see anything on the SR that I opened to Check Point.
Hope it helps
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